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What the Fathers say
Western Saints
Augustine of Hippo
What then, when the sun went down, when our Lord suffered? There was a sort of darkness with the Apostles, hope failed, in those to whom He at first seemed great, and the Redeemer of all men. How so? "Thou didst make darkness, and it became night; wherein all the beasts of the forest shall move" (ver. 20). ...Here the beasts of the forest are used in different ways: for these things are always understood in varying senses; as our Lord Himself is at one time termed a lion, at another a lamb. What is so different as a lion and a lamb? …
Jerome
You have set darkness, and it became night: in it all the beasts of the forest shall pass through. And according to the letter it has its own order: for night was made for this reason, that men might rest from their works, and that the beasts might go out to their feeding. But according to the figurative interpretation, when the sun of righteousness has set upon us, all things are dark to us, and then the beasts pass through against us, and the roaring lion, wishing to seize and devour us.
Translation